With the bulk modulus and density of the seagrass tissue determined by previous work, this study focused on characterizing the shear moduli of the epidermis and aerenchyma. For this purpose, a new seagrass leaf model is described for Thalassia testudinum that consists of a comparatively rigid epidermis that composes the outer shell of the leaf and comparatively compliant aerenchyma that surrounds the gas channels on the interior of the leaf. To make accurate assessments of seagrass biomass and productivity, a model is needed to describe acoustic propagation through the seagrass meadow that includes the effects of gas contained within the seagrass leaves. Acoustic monitoring techniques exploit the sensitivity of underwater sound to bubbles, which are produced as a byproduct of photosynthesis and present within the seagrass tissue. Seagrasses play an important role in coastal ecosystems and serve as important marine carbon stores.
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